Notholaena
| Notholaena | |
|---|---|
 | |
| N. standleyi | |
| Scientific classification | |
| Kingdom: | Plantae |
| Division: | Pteridophyta |
| Class: | Polypodiopsida |
| Order: | Polypodiales |
| Family: | Pteridaceae |
| Subfamily: | Cheilanthoideae |
| Genus: | Notholaena R.Br. |
| Species | |
|
See text. | |
Notholaena, cloak fern,[1] is a genus of ferns in the Cheilanthoideae subfamily of the Pteridaceae.[2] Ferns of this genus are mostly epipetric (growing on rock) or occurring in coarse, gravelly soils, and are most abundant and diverse in the mountain ranges of warm arid or semiarid regions. They typically have a creeping or erect rhizome and leaves that are pinnatifid to pinnate-pinnatifid with marginal sori protected by a false indusium formed from the reflexed margin of the leaf. Members of Notholaena also have a coating of whitish or yellowish farina (a powdery wax that prevents desiccation) on the surfaces of the leaves. The farina is often limited to the abaxial (lower) leaf surface, but may occur on the adaxial (upper) leaf surface as well.
The similar genus Argyrochosma also has farinose leaves, but in that genus the ultimate segments of the leaves have entire margins and *are distinctly stalked, whereas in Notholaena the ultimate segments are usually lobed or pinnatifid and sessile or subsessile. Notholaena has in the past been used as a "catch-all" genus for a wide variety of species that did not fit well in other arid fern genera but it has more recently been defined in a much narrower sense, making the genus much more morphologically and evolutionarily coherent. The genera Argyrochosma and Astrolepis were recently segregated from Notholaena, and other former members of Notholaena are now in the genus Cheilanthes.
Selected species
- Notholaena bryopoda – a species unusual among ferns in that it is endemic to gypseous substrates in Mexico
- Notholaena californica – a fern of the southwestern US and northern Mexico
- Notholaena grayi – a fern of igneous substrates in Mexico and the Madrean sky islands of the southwestern US
- Notholaena marantae – occurs on dry rocks and serpentines in South and Central Europe - it is now placed in the genus Paragymnopteris
- Notholaena standleyi – a common fern of igneous substrates in the southwestern US
References
- ↑ "Notholaena". Natural Resources Conservation Service PLANTS Database. USDA. Retrieved 20 July 2015.
- ↑ Christenhusz, Maarten J. M.; Zhang, Xian-Chun; Schneider, Harald (18 February 2011). "A linear sequence of extant families and genera of lycophytes and ferns" (PDF). Phytotaxa. 19: 7–54. ISSN 1179-3163.