List of archosaurs of the Chinle Formation

The Chinle Formation is an Upper Triassic continental geologic formation in the western United States which has yielded fossils of many archosaurs: a group of vertebrates that includes crocodiles, pterosaurs, dinosaurs (including birds), and other extinct relatives.

Crurotarsans

Aetosaurs

Color key

Taxon	Reclassified taxon	Taxon falsely reported as present	Dubious taxon or junior synonym	Ichnotaxon	Ootaxon	Morphotaxon

Notes
Uncertain or tentative taxa are in small text; ~~crossed out~~ taxa are discredited.

Misc. Archosauromorphs of the Chinle Formation
Genus	Species	State	Member	Abundance	Notes	Images
Acaenasuchus^[1]	A. geoffreyi^[1]	Arizona^[1]	Sonsela^[1] Blue Mesa^[1] Mesa Redondo^[1]	Known from a handful of isolated osteoderms.^[1]	A controversial taxon that some paleontologists have considered to be juvenile Desmatosuchus haplocerus scutes.^[1] However, Randall Irmis notes that the Acaenasuchus scutes have unique characters.^[1] Some earlier workers attributed these characters to the allegedly young age of the specimens.^[1] However, Irmis notes that the ontogenetic stage of the specimens has not been confirmed and that in aetosaur genera with known juvenile remains, there are no appreciable physical differences between the younger and older individuals.^[1] Irmis suggested that future researchers treat Acaenasuchus as its own taxon until contrary evidence surfaces.^[1]	Desmatosuchus Stagonolepis
Desmatosuchus^[2]	D. spurensis^[3]	Arizona^[3]	Sonsela^[3] Blue Mesa^[3] Mesa Redondo^[3]
Desmatosuchus^[2]	D. smalli^[4]	Arizona^[4]	Petrified Forest^[4]
Rioarribasuchus	R. chamaensis	Arizona^[5] New Mexico^[5]	Petrified Forest^[5]		Although originally referred to Desmatosuchus, Randall Irmis considers it to be a new taxon closely related to Paratypothorax based on a phylogenetic analysis performed on the remains by Parker in 2003.^[5]
Paratypothorax^[6]	P. andressorum^[6]	Arizona^[6]			A significant number of researchers feel that the North American material referred to the German Paratypothorax species P. andressorum by Hunt and Lucas in 1992 is actually distinct.^[6]
Paratypothorax^[6]	Indeterminate^[7]	Arizona^[7]	Sonsela^[7]
Calyptosuchus^[8]	C. wellesi^[9]	Arizona^[9]	Sonsela^[9] Blue Mesa^[9] Mesa Redondo^[9]	Common in the lower strata of the Chinle Formation, although "[n]on-armor material is comparatively rare".^[9]
Calyptosuchus^[8]	Indeterminate^[10]	Arizona^[10]	Sonsela^[10]		A partial skeleton from Petrified Forest National Park is not only the uppermost occurrence of Stagonolepis in the park, but may represent a new species as well.^[10]
Typothorax^[11]	T. coccinarum^[11]	Arizona^[11]	Owl Rock^[11] Petrified Forest^[11] Sonsela^[11]	Common in the Owl Rock and Petrified Forest Members.^[11]	Almost all of the known Chinlean Typothorax fossils are isolated whole or fragmentary osteoderms.^[11]

Crocodylomorphs

Rauisuchians of the Chinle Formation
Genus	Species	State	Member	Abundance	Notes	Images
Hesperosuchus^[12]	H. agilis^[12]	Arizona^[12]	Petrified Forest^[12] Blue Mesa^[12] Mesa Redondo^[12]			Hesperosuchus
Parrishia^[13]	P. mccreai^[13]	Arizona^[13]	Blue Mesa^[13] Mesa Redondo^[13]

Phytosaurs

Phytosaurs of the Chinle Formation
Genus	Species	State	Member	Abundance	Notes	Images
Leptosuchus^[14]	L. crosbiensis^[14]	Arizona^[14]	Sonsela^[14] Blue Mesa^[14] Mesa Redondo^[14]
Smilosuchus^[15]	S. adamanensis^[14]	Arizona^[14]	Sonsela^[14] Blue Mesa^[14] Mesa Redondo^[14]	Known from a single skull	Previously referred to Leptosuchus, actually referable to Smilosuchus.^[16]
	S. gregorii	Arizona	Sonsela Blue Mesa Mesa Redondo	Known from a handful of skulls.	A basal leptosuchomorph.	Smilosuchus
	S. lithodendrorum^[16]	Arizona	Blue Mesa	Known from only one skull.	Previously considered synonymous with S. adamanensis.
cf. Paleorhinus^[17]	Indeterminate^[17]	Arizona^[17]	Mesa Redondo^[17]
Machaeroprosopus^[18]	M. buceros^[18]	Arizona^[18]	Owl Rock^[18] Petrified Forest^[18] Sonsela^[18]
	M. mccauleyi^[18]	Arizona^[18]	Owl Rock^[18] Petrified Forest^[18] Sonsela^[18]
	M. pristinus^[18]	Arizona^[18]	Owl Rock^[18] Petrified Forest^[18] Sonsela^[18]
Redondasaurus^[19]	R. gregorii^[19]	Arizona^[15] Utah^[15]	Rock Point^[15] Petrified Forest^[15] Church Rock^[15]		Long and Murray considered Redondasaurus to be synonymous with Pseudopalatus,^[15] but other workers have found in to be distinct.^[20]

Rauisuchians

Rauisuchians of the Chinle Formation
Genus	Species	State	Member	Abundance	Notes	Images
Shuvosaurus^[21]	S. inexpectatus^[21]	Arizona^[21]	Owl Rock^[21] Petrified Forest^[21] Mesa Redondo^[21]			Effigia Poposaurus Postosuchus Shuvosaurus
Effigia	E. okeeffeae	New Mexico	Ghost Ranch
Poposaurus^[22]	P. gracilis^[22]	Arizona^[22]	Blue Mesa^[22] Mesa Redondo^[22]
Postosuchus^[23]	P. kirkpatricki^[23]	Arizona^[23]	Owl Rock^[23] Petrified Forest^[23] Sonsela^[23] Blue Mesa^[23] Mesa Redondo^[23]
Saurosuchus^[24]	Indeterminate^[24]	Arizona^[24]	Blue Mesa^[24]
Vivaron^[25]	V. haydeni^[25]	New Mexico^[25]	Petrified Forest^[25]

Other crurotarsans

Misc. Crurotarsans of the Chinle Formation
Genus	Species	State	Member	Abundance	Notes
Revueltosaurus^[26]	R. callenderi^[27]	Arizona^[27]	Petrified Forest^[27]
Revueltosaurus^[26]	R. hunti^[28]	Arizona^[28]	Blue Mesa^[28]

Dinosauriformes

Basal dinosauriforms

Color key

Taxon	Reclassified taxon	Taxon falsely reported as present	Dubious taxon or junior synonym	Ichnotaxon	Ootaxon	Morphotaxon

Notes
Uncertain or tentative taxa are in small text; ~~crossed out~~ taxa are discredited.

Basal Dinosauriforms of the Chinle Formation
Genus	Species	State	Member	Abundance	Notes	Images
Eucoelophysis	E. baldwini	New Mexico^[29]	Petrified Forest^[29]		A dinosauriform closely related to the Polish genus Silesaurus.^[30]

Dinosaurs

Saurischians

Sauropodomorphs

Prosauropod tracks are present in the Redonda, Sloan Canyon, and Sheep Pen Sandstone formations. Possibly the Rock Point Formation as well. Geographically, the tracks are present in New Mexico.^[29]

Theropods

Indeterminate ceratosaurian remains have been recovered from the Bull Canyon Formation in New Mexico.^[29] Theropod tracks have been found in Utah and New Mexico recovered from the Redonda, Sloan Canyon, and Sheep Pen Sandstone formations.^[29] Indeterminate theropod remains are stratigraphically present in the Santa Rosa, Petrified Forest, Bluewater Creek, Rock Point, and Garita Creek formations. They are geographically present in New Mexico.^[29] Indeterminate herrerasaurid remains stratigraphically present in the Bull Canyon Formation and geographically present in New Mexico.^[29]

Theropods of the Chinle Formation
Genus	Species	State	Member	Abundance	Notes	Images
Camposaurus^[29]	C. arizonensis^[29]	Arizona^[29]	Bluewater Creek Formation^[29]		A coelophysine coelophysid	Camposaurus Chindesaurus Coelophysis Gojirasaurus Tawa
Chindesaurus^[29]	C. bryansmalli^[29]	Arizona^[29] New Mexico^[29]	Bull Canyon Formation^[29] Petrified Forest^[29]		A herrerasaur
Coelophysis^[29]	C. bauri	New Mexico^[31]	Siltstone^[31]
	C. sp.^[32]	Arizona^[32]	Petrified Forest^[32]		Originally assigned to C. bauri, is likely a different taxon.^[32]
	C. longicollis^[29]	New Mexico^[29]	Petrified Forest^[29]		Now considered a junior synonym of C. bauri.
	C. willistoni	New Mexico^[29]	Petrified Forest^[29]		Now considered a junior synonym of C. bauri.
Daemonosaurus	D. chauliodus	New Mexico	"Siltstone"		A basal theropod
Gojirasaurus^[29]	G. quayi^[29]	New Mexico^[29]	Cooper Canyon^[29]		A coelophysoid
Tawa	T. hallae	New Mexico			A basal theropod

Footnotes

1 2 3 4 5 6 7 8 9 10 11 12 "Vertebrate Fauna; Stagonolepididae; Acaenasuchus geoffreyi," Irmis (2005) pp. 75-77
↑ "Vertebrate Fauna; Stagonolepididae," Irmis (2005) p. 75
1 2 3 4 5 "Vertebrate Fauna; Stagonolepididae; Desmatosuchus haplocerus," Irmis (2005) p. 75
1 2 3 "Vertebrate Fauna; Stagonolepididae; Desmatosuchus smalli," Irmis (2005) p. 75
1 2 3 4 "Vertebrate Fauna; Stagonolepididae; "Desmatosuchus" chamaensis," Irmis (2005) p. 75
1 2 3 4 "Vertebrate Fauna; Stagonolepididae; Paratypothorax sp.," Irmis (2005) p. 78
1 2 3 "Vertebrate Fauna; Stagonolepididae; 'Paratypothorax sp.' and 'Paratypothorax-Like Forms'," Irmis (2005) p. 78
↑ "Vertebrate Fauna; Stagonolepididae," Irmis (2005) p. 77
1 2 3 4 5 6 "Vertebrate Fauna; Stagonolepididae; Stagonolepis wellesi," Irmis (2005) p. 77
1 2 3 4 "Vertebrate Fauna; Stagonolepididae; Stagonolepis sp." Irmis (2005) p. 77
1 2 3 4 5 6 7 8 "Vertebrate Fauna; Stagonolepididae; Typothorax coccinarum," Irmis (2005) pp. 77-78
1 2 3 4 5 6 "Vertebrate Fauna; Crocodylomorpha; Hesperosuchus agilis," Irmis (2005) pp. 79-80
1 2 3 4 5 "Vertebrate Fauna; Crocodylomorpha; 'Parrishia mccreai,'" Irmis (2005) pp. 80-81
1 2 3 4 5 6 7 8 9 10 11 "Vertebrate Fauna; Parasuchia; Leptosuchus adamanensis, L. crosbiensis, and L. gregorii," Irmis (2005) p. 72, 74
1 2 3 4 5 6 7 Long, R.A. and Murray, P.A. 1995. Late Triassic (Carnian and Norian) Tetrapods from the Southwestern United States. New Mexico Museum of Natural History and Science Bulletin 4. pp. 254.
1 2 Michelle R. Stocker (2010). "A new taxon of phytosaur (Archosauria: Pseudosuchia) from the Late Triassic (Norian) Sonsela Member (Chinle Formation) in Arizona, and a critical reevaluation of Leptosuchus Case, 1922". Palaeontology 53 (5): 997–1022. doi:10.1111/j.1475-4983.2010.00983.x.
1 2 3 4 "Vertebrate Fauna; Parasuchia; 'Paleorhinus sp.'" Irmis (2005) p. 72
1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 "Vertebrate Fauna; Parasuchia; Pseudopalatus pristinus, P. buceros, and P. mccauleyi," Irmis (2005) p. 74-75
1 2 Hunt, A.P. and Lucas, S.G. 1993. A new phytosaur (Reptilia: Archosauria) genus from the uppermost Triassic of the western United States and its biochronological significance. In; The Nonmarine Triassic, Lucas and Morales, eds., New Mexico Museum of Natural History and Science Bulletin 3.
↑ Hungerbühler A. 2002. The Late Triassic phytosaur Mystriosuchus westphali, with a revision of the genus. Palaeontology 45 (2): 377-418
1 2 3 4 5 6 "Vertebrate Fauna; Poposauridae; 'Chatterjeea elegans,'" Irmis (2005) p. 79
1 2 3 4 5 "Vertebrate Fauna; Poposauridae; Poposaurus gracilis," Irmis (2005) p. 79
1 2 3 4 5 6 7 8 "Vertebrate Fauna; 'Rauisuchidae'; Postosuchus kirkpatricki," Irmis (2005) p. 78
1 2 3 4 "Vertebrate Fauna; 'Rauisuchidae'; 'Saurosuchus sp.,'" Irmis (2005) p. 78
1 2 3 4 Emily J. Lessner; Michelle R. Stocker; Nathan D. Smith; Alan H. Turner; Randall B. Irmis; Sterling J. Nesbitt (2016). "A new rauisuchid (Archosauria, Pseudosuchia) from the Upper Triassic (Norian) of New Mexico increases the diversity and temporal range of the clade". PeerJ. 4: e2336. doi:10.7717/peerj.2336.
↑ "Vertebrate Fauna; Pseudosuchia; 'Revueltosaurus callenderi' and 'Revueltosaurus hunti,'" Irmis (2005) pp. 71-72
1 2 3 "Vertebrate Fauna; Pseudosuchia; Revueltosaurus callenderi," Irmis (2005) pp. 71-72
1 2 3 "Vertebrate Fauna; Pseudosuchia; Revueltosaurus hunti," Irmis (2005) p. 72
1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 Weishampel, David B; et al. (2004). "Dinosaur distribution (Late Triassic, North America)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 518–521. ISBN 0-520-24209-2.
↑ See Nesbitt et al. (2005) and Ezcurra (2006).
1 2 Zeigler, K. E.; Kelley, S.; Geissman, J. W. (2008). "Revisions to stratigraphic nomenclature of the Upper Triassic Chinle Group in New Mexico: New insights from geologic mapping, sedimentology, and magnetostratigraphic/paleomagnetic data". Rocky Mountain Geology. 43 (2): 121–141. doi:10.2113/gsrocky.43.2.121.
1 2 3 4 "Vertebrate Fauna; Pseudosuchia; Revueltosaurus callenderi," Irmis (2005) p. 81

References

Ezcurra, M.D. (2006). "A review of the systematic position of the dinosauriform archosaur Eucoelophysis baldwini Sullivan & Lucas, 1999 from the Upper Triassic of New Mexico, USA." Geodiversitas, 28(4):649-684.
Irmis, R. B. 2005. The vertebrate fauna of the Upper Triassic Chinle Formation in northern Arizona. p. 63-88. in S.J. Nesbitt, W.G. Parker, and R.B. Irmis (eds.) 2005. Guidebook to the Triassic formations of the Colorado Plateau in northern Arizona: Geology, Paleontology, and History. Mesa Southwest Museum Bulletin 9.
Nesbitt, S.J., Irmis, R.B., and Parker, W.G. (2005). "A critical review of the Triassic North American dinosaur record." In Kellner, A.W.A., Henriques, D.D.R., & Rodrigues, T. (eds.), II Congresso Latino-Americano de Paleontologia de Vertebrados, Boletim de Resumos. Rio de Janeiro: Museum Nacional/UFRJ, 139.
Sues, H.-D., Nesbitt, S. J., Berman, D. S., and A. C., Henrici. 2011. A late-surviving basal theropod dinosaur from the latest Triassic of North America. Proceedings of the Royal Society B. doi:10.1098/rspb.2011.0410
Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. 861 pp. ISBN 0-520-24209-2.

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